Chytridiomycota is a division of zoosporic organisms in the kingdom Fungiinformally known as chytrids. Chytrids are saprobicdegrading refractory materials such as chitin and keratinand sometimes act as parasites.
Species of Chytridiomycota have traditionally been delineated and classified based on development, morphology, substrate, and method of zoospore discharge.
In an older and more restricted sense not used herethe term "chytrids" referred just to those fungi in the class Chytridiomycetes. The chytrids have also been included among the Protoctista but are now regularly as fungi.
In older classificationschytrids, except Chytridiomycota asexual reproduction in fungi recently established order Spizellomycetaleswere placed in the class Phycomycetes under the subphylum Myxomycophyta of the kingdom Fungi. Previously, they were placed in the Mastigomycotina as the class Chytridiomycetes.
The class Chytridiomycetes has over chytrid species distributed among ten orders.
Molecular phylogeneticsand other techniques such as ultrastructure analysis, has greatly increased the understanding of chytrid phylogeny, and led to the formation of several new zoosporic fungal phyla:. Chytridiomycota are unusual among the Fungi in that they reproduce with zoospores. Asexual reproduction occurs through the release of zoospores presumably derived through mitosis.
Where it has been described, sexual reproduction of Chytridomycetes occurs via a variety of methods. It is generally accepted that the resulting zygote forms a resting spore, which functions as a means of surviving adverse conditions.
This group Chytridiomycota asexual reproduction in fungi the notable plant pathogens Synchytrium. Some algal parasites practice oogamy: In another group, two thalli produce tubes that fuse and allow the gametes to meet fuse.
Both nuclei migrate out of the zoosporangium and into the conjoined rhizoids where they fuse. The resulting zygote germinates into a resting spore. Sexual reproduction is common and well known among members of the Monblepharidomycetes.
Typically, these chytrids practice a version of oogamy: This is the first occurrence of oogamy in kingdom Fungi. Once fertilized, the zygote either becomes an encysted or motile oospore,  which ultimately becomes a resting spore that will later and give rise to new zoosporangia.
Upon release from the germinated resting spore, zoospores seek out
Chytridiomycota asexual reproduction in fungi suitable substrate for growth using chemotaxis or
Chytridiomycota asexual reproduction in fungi. Some
Chytridiomycota asexual reproduction in fungi encyst and germinate directly upon the substrate; others encyst and germinate a short distance Chytridiomycota asexual reproduction in fungi. Once germinated, enzymes released from the zoospore begin to break down the substrate and utilize it produce a Chytridiomycota asexual reproduction in fungi thallus.
Thalli are coenocytic and usually form no true mycelium having rhizoids instead. Chytrids have several different growth patterns. Some are Chytridiomycota asexual reproduction in fungi, which means they only produce a zoosporangium and zoospores. Others are eucarpic, meaning they produce other structures, such as rhizoidsin addition to the zoosporangium and zoospores. Some chytrids are monocentric,
Chytridiomycota asexual reproduction in fungi a single zoospore gives rise to a single zoosporangium.
Others are polycentric, meaning one zoospore gives rise to many zoosporangium connected by a rhizomycelium. Rhizoids do not have nuclei while a rhizomycelium can. Growth continues until a new batch of zoospores are ready for release. Chytrids have a diverse set of release mechanisms that can be grouped into the broad categories of operculate or inoperculate.
Operculate discharge involves the complete or incomplete detachment of a lid-like structure, called an operculum, allowing the zoospores out of the sporangium. Inoperculate chytrids release their zoospores through pores, slits, or papillae. Chytrids are aquatic fungithough those that thrive in the capillary network around soil particles are typically considered terrestrial. Chytrids have been isolated from a variety of aquatic habitats, including peats, bogs, rivers, ponds, springs, and ditches, and terrestrial habitats, such as acidic soils, alkaline soils, temperate forest soils, rainforest soils, Arctic and Antarctic soils.
One of the least expected terrestrial environments the chytrid thrive in are periglacial soils. The chytrid Batrachochytrium dendrobatidis is responsible for chytridiomycosisa disease of amphibians. Discovered in in Australia and Panama this disease is known to kill amphibians in large numbers, and has been suggested as a principal cause for the worldwide amphibian decline.
Outbreaks of the fungus were found responsible for killing much of the Kihansi Spray Toad population Chytridiomycota asexual reproduction in fungi its native Chytridiomycota asexual reproduction in fungi of Tanzania as well as the extinction of the golden toad in Chytridiomycosis has also implicated in the presumed extinction of the Southern Gastric Brooding Frog last seen in the wild inand the Northern Gastric Brooding Frog, last recorded in the wild in March .
The process leading to frog mortality is thought to be the loss of essential ions through pores made in the epidermal cells by the chytrid during its replication. Recent research has revealed that elevating salt levels slightly may be able to cure chytridiomycosis in some Australian frog species  although further Chytridiomycota asexual reproduction in fungi is needed. Chytrids mainly infect algae and other eukaryotic and prokaryotic microbes.
The infection can be so severe as to control primary production within the lake. Arguably, the most important ecological function chytrids perform is decomposition. This colonization Chytridiomycota asexual reproduction in fungi pollen happens during the spring time when bodies of water accumulate pollen falling from trees and plants. Among the microfossils are chytrids preserved as parasites on rhyniophytes. These
Chytridiomycota asexual reproduction in fungi closely resemble the modern genus Allomyces.
These remains were found along with eucarpic remains and are ambiguous in nature although they are thought to be of chytrids. The novel Tom Clancy's Splinter Cell: Fallout features a species of chytrid that feeds on petroleum and oil-based products.
In the story the species is modified using nuclear radiationto increase the rate at which it feeds on oil. It is then used by Islamic extremists in an attempt to destroy the world's oil supplies, thereby taking away the technological advantage of the United States. The webcomic Black Mudpuppy  by Ethan Kocak features an arc  with an evil, anthropomorphic chytrid developed
Chytridiomycota asexual reproduction in fungi a Nazi experiment to infect Xolotl, a giant, talking black axolotl.
From Wikipedia, the free encyclopedia. Chytridiomycota Sporangium of a spizellomycete Scientific classification Kingdom: Chytridiomycota Hibbett et al. The University of Michigan Press: American Journal of Botany Thallus development and the systematics of Chytridiomycota: Basidiomycetes and Lower Archived from the original on Systematics and Evolution 2nd ed.
A new genus and family for Chytridiomycota asexual reproduction in fungi misclassified chytrid, Rhizophlyctis harderi in press. Deshmukh 10 January Systematics and Evolution of Fungi. A higher-level phylogenetic classification of the Fungi. The zoospore and its problems. Water, Fungi, and Plants. Cambridge University Press, Rhizophlyctidales--a new order in Chytridiomycota.
Phylogeny of Powellomycetacea fam. Temporal variation of the small eukaryotic community in two freshwater lakes: Aquatic Microbial Ecology Retrieved 28 October The New York Times.
Electrolyte depletion and osmotic imbalance in amphibians with chytridiomycosis. Host parasite interactions between freshwater phytolankton and chytrid fungi Chytridiomycota asexual reproduction in fungi. Compendium of Potato Diseases. Chytridiomycota and chytrid-like remains of uncertain affinity". Review of Palaeobotany and Palynology. Taylor; Nora Dotzler Tom Clancy's Splinter Cell: True fungi classification, fungal orders.
Xylonomycetes Geoglossomycetes Leotiomycetes Chytridiomycota asexual reproduction in fungi Sordariomycetes. Archaeorhizomycetes Neolectomycetes Pneumocystidomycetes Schizosaccharomycetes Taphrinomycetes. Tritirachiomycetes Mixiomycetes Agaricostilbomycetes Cystobasidiomycetes Microbotryomycetes Classiculomycetes Cryptomycocolacomycetes Atractiellomycetes Pucciniomycetes. Monilielliomycetes Malasseziomycetes Ustilaginomycetes Exobasidiomycetes. Hyaloraphidiomycetes Monoblepharidomycetes Sanchytriomycetes Mesochytriomycetes Chytridiomycetes.
Fungal phyla are underlined. Retrieved from " https: Chytridiomycota Aquatic fungi Fungus phyla Fungi by classification. Archived copy as title with 'species' Chytridiomycota asexual reproduction in fungi Commons category link is on Wikidata. Views Read Edit View history. In other projects Wikimedia Commons Wikispecies. This page was last edited on 24 Octoberat